The Model 290 UHF data link transponder is a command receiver, digital command decoder, digital telemetry encoder, telemetry transmitter, and target interface enclosed in a single lightweight package. The transponder receives commands transmitted by the ground control station and interfaces with the vehicle management computer or autopilot.

Uplink UHF command messages from the ground control station are received by the transponder in the frequency band of 380 MHz to 435 MHz. The RF data is converted to a serial data stream that is digitally transmitted to the vehicle autopilot for processing. The autopilot can then send a serial data stream back to the transponder, where the telemetry downlink is then transmitted down to the ground control station. It also has three antenna ports to support multiple antenna inputs via the vehicle RF switch. Multiple antenna inputs can be toggled through for the strongest signal.

The Model 290 has been qualified per MIL-STD-810 (Environmental), making it one of the most capable devices in the industry.